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MATERIALS
Recent and fossil specimens examined for this study belong to: Geological Institute of the Siberian Branch RAS, Ulan-Ude; Institute of Plant and Animal Ecology USC, Yekaterinburg; Zoological Institute RAS, Saint Petersburg, Russia and Institute of Geology SB RAS, Yakutsk (Russia); American Museum of Natural History, New York; Smithsonian Institution, Washington, D.C.; Carnegie Museum of Natural History, Pittsburgh; Denver Museum of Nature and Science; and the University of California, Museum of Paleontology (USA).
 
The systematic organization of fossil species presented below is based on the morphology of preserved skulls, mandibles and teeth, and on biometric studies. Most living species, in addition to the mentioned characters, were diagnosed according to external features such as body ratio or pelage. The comparative analysis of living and extinct species is based mainly on the tooth structures and in part on preserved fragments of skull and mandibles for fossil forms.
 
We follow the standard Ochotona dental morphology outlined by 
Erbajeva (1988). In a whole more than 2000 specimens of living pikas and approximately 650 specimens of fossil Ochotona were examined from mentioned above Institutions.
 
Classification of the family Ochotonidae
Five families are recognized in the order Lagomorpha: Mimolagidae
Erbajeva, 1986, Leporidae
Gray, 1821, Palaeolagidae
Dice, 1929, Prolagidae
Gureev, 1960, and Ochotonidae
Thomas, 1897.  The family Ochotonidae includes two subfamilies, Sinolagomyinae
Gureev, 1960 and Ochotoninae
Thomas, 1897. The chronologic range of Sinolagomyinae is Oligocene to Miocene.  Ochotoninae is known from the Miocene – Recent.  The classification of genera (after 
Erbajeva 1994, modified with new data from
Sen, 2003) is as follows:
 
Ochotonidae Thomas, 1897
 
Sinolagomyinae Gureev, 1960
 
  
Sinolagomys Bohlin, 1937  
Austrolagomys Stromer, 1926  
Kenyalagomys MacInnes, 1953  
Oreolagus Dice, 1917  
Bellatona Dawson, 1961  
Heterolagus 
Crusafont, Villata, and Truyols, 1955  
Bellatonoides Sen, 2003
 
   
Ochotoninae Thomas, 1897
 
  
Marcuinomys Groizet, 1839  
Lagopsis Schlosser, 1884  
Albertona Lopez-Martinez, 1986  
Alloptox Dawson, 1961  
Paludotona Dawson, 1959  
Proochotona Chomenko, 1914  
Ochotonoides
Teilhard de Chardin and Young, 1931  
Pliolagomys 
Erbajeva, 1983 
in Agadjanian and Erbajeva, 1983 
Ochotonoma Sen, 1998  
Ochotona Link, 1795
 
   
 
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